Foliage simulation

A foliage simulation based on a disney research paper (link in below).
The system uses sunlight, soil fertility, local competition, temperature and elevation to simulate an environment.
In the video you can see a simulation of 76 years (one year per frame stretched out) in a maritime climate.

(research paper: https://s3-us-west-1.amazonaws.com/disneyresearch/wp-content/uploads/20160608160305/Guided-Ecological-Simulation-for-Artistic-Editing-of-Plant-Distributions-in-Natural-Scenes-Paper.pdf)